Dr. David Lees from the Department of Zoology of the University of Cambridge, United Kingdom, and the Natural History Museum in London, helped me identifying this species. 
I met David in Madagascar in January 2014 during his research work in Andasibe-Mantadia NP and had the pleasure to spent a few days together.
David has extensive field research experience in Madagascar and is considered the leading expert on Madagascar butterflies.

Nosy Komba, Madagascar, 24 February 2020