Dr. David Lees from the Department of Zoology of the University of Cambridge, United Kingdom, and the Natural History Museum in London, helped me identify this species. 
I met David in Madagascar in January 2014 during his research work in Andasibe-Mantadia NP and had the pleasure of spending a few days together.
David has extensive field research experience in Madagascar and is considered the leading expert on Madagascar butterflies.

Enjojo Lodge, Ishasha area, Queen Elizabeth National Park, Uganda, 21 January 2025